Resident Evil 5 features similar gameplay to Resident Evil 4. Context-sensitive controls and dynamic cut scenes return in Resident Evil 5.

The player(s) control Chris Redfield and/or Sheva Alomar, similarly to how the player controlled Leon S. Kennedy in Resident Evil 4, with the same over-the-shoulder perspective. The game's environment plays a significant role. The Mercenaries minigame, which was featured in both Resident Evil 3 and Resident Evil 4, is also in Resident Evil 5. As of launch, the multiplayer in the mini-game was offline only, but a launch day patch gave the game online multi-player.
The game features new types of enemies called "Majini", meaning evil spirit in Swahili. The game's enemies possess human-like intelligence and awareness. Furthermore, they are different from the previous ganados. The number of weapon variations has been greatly increased compared to the previous games in the series. Requisites such as handguns and shotguns also return.

The game features some online elements. Resident Evil 5's main story mode can be played with two players, in a co-operative online mode. The feature will allow players to enter or leave anytime during the game. Players will not always stick together, and can be separated at points during the gameplay. An offline co-op mode is also in the game with the same co-op experience offered by the online co-op mode, only with two local players controlling the action in split screen.

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